Botanist and plant pathologist Effie (Southworth) Spalding measuring a saguaro rib. An accomplished scientist, Effie worked at the Desert Botanical Laboratory of the Carnegie Institute (known today as Tumamoc: People & Habitats) from 1905-1911. She published the very first research paper done on Tumamoc in 1905. Her research showed that the ribs of saguaro cacti expand and contract as they store and use water.
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